Her passing wasn't a complete surprise, zoo officials said. She'd been ill in recent months.
April arrived at the Minnesota Zoo in January 2008, on loan from the Brookfield Zoo in Chicago along with her daughter, Allie. Last summer, April became a grandma when Allie gave birth to Taijah.
"April was one of our most beloved animals," zoo director of biological programs Kevin Willis said. "Since she arrived here a few years ago, she quickly became a guest favorite. She enjoyed interacting with her trainers during enrichment sessions, and was often heard vocalizing during these interactions. She will be terribly missed by staff, volunteers, and all the zoo guests who met her."
Here's April's granddaughter, Taijah, swimming with her father, Semo: